Animals Defend Their Territory

Behavioural ecologists have argued that food distribution determines whether a species will be territorial or not. Territoriality will emerge where there is a focused resource that provides enough for the individual or group, within a boundary that is small enough to be defended without the expenditure too much effort.
Territoriality is least likely with insectivorous birds, where the food supply is plentiful but unpredictably distributed. Swifts rarely defend an area larger than the nest.
Conversely, large solitary (or paired) carnivores, such as bears and the bigger raptors require an extensive protected area to guarantee their food supply. This territoriality will only break down when there is a glut food, for example when Grizzly Bears are attracted to migrating salmon.