Emotion In Animals

Primates and in particular great apes are candidates for highly developed capabilities for empathy and theories of mind. Great apes have highly complex social systems. Young apes and their mothers have very strong bonds of attachment. Often when a baby chimpanzee or gorilla dies, the mother will carry the body around for several days. Jane Goodall has described chimpanzees as exhibiting mournful behavior. Notably the example of the gorilla Koko, who expressed sadness over the death of her pet cat, All Ball.
Canines
Research suggests that canines can experience negative emotions in a similar manner to people, including the equivalent of certain chronic and acute psychological conditions. The classic experiment for this was Martin Seligman's foundational experiments and theory of learned helplessness at the University of Pennsylvania in 1965, as an extension of his interest in depression: