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Insect Macro Photography
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Insects were among the earliest terrestrial herbivores and acted as major selection agents on plants. Plants evolved chemical defenses against this herbivory and the insects in turn evolved mechanisms to deal with plant toxins. Many insects make use of these toxins to protect themselves from their predators. Such insects often advertise their toxicity using warning colors. This successful evolutionary pattern has also been utilized by mimics. Over time, this has led to complex groups of coevolved species. Conversely, some interactions between plants and insects, like pollination, are beneficial to both organisms. Coevolution has led to the development of very specific mutualisms in such systems.
• Taxonomy
Traditional morphology-based or appearance-based systematics has usually given Hexapoda the rank of superclass, and identified four groups within it: insects (Ectognatha), springtails (Collembola), Protura and Diplura, the latter three being grouped together as Entognatha on the basis of internalized mouth parts. Supraordinal relationships have undergone numerous changes with the advent of methods based on evolutionary history and genetic data. A recent theory is that Hexapoda is polyphyletic (where the last common ancestor was not a member of the group), with the entognath classes having separate evolutionary histories from Insecta. Many of the traditional appearance-based taxa have been shown to be paraphyletic, so rather than using ranks like subclass, superorder and infraorder, it has proved better to use monophyletic groupings (in which the last common ancestor is a member of the group). The following represents the best supported monophyletic groupings for the Insecta.
Insects can be divided into two groups historically treated as subclasses: wingless insects, known as Apterygota, and winged insects, known as Pterygota. The Apterygota consist of the primitively wingless order of the silverfish (Thysanura). Archaeognatha make up the Monocondylia based on the shape of their mandibles, while Thysanura and Pterygota are grouped together as Dicondylia. It is possible that the Thysanura themselves are not monophyletic, with the family Lepidotrichidae being a sister group to the Dicondylia (Pterygota and the remaining Thysanura).
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